Plant Guide to Aspen Grove / Fish Creek Trail, San Bernardino Mountains

This is a working list, about which we make no guarantees at all until we officially release it. Use at your own risk!

See also Plant Voucher Checklist for Fish Creek Area for a plant list in traditional family order, referenced to this guide, the guide to the Upper Fish Creek Trail, and including vouchers.

Introduction and Explanation of Plant Trail Guides

Highlights of This Trail
Fieldwork Dates and Summary of List Changes With Time
Botanical Trip Reports
The Plant Guide
Comments On Specific Species


Directions to trailhead: On SR38, go 5 miles east of Barton Flats to the signed entrance road to Heart Bar Campground, 1N02. Do not take the earlier turnoff to Camp Heart Bar! Turn right on 1N02 to a signed junction in 1.2 miles with 1N05. Go right, up 1N05, bearing right at all road forks.

The signed Aspen Grove / Lower Fish Creek Trailhead parking is reached a total distance of 2.6 miles from SR38, which takes about 10 minutes total driving time.

GPS Coordinates:

TOPO! GPS Data Format Deg NAD27 ElevFeet

1N05 is the junction of 1N02 and 1N05.
FISHLO is the Aspen Grove / Lower Fish Creek Trailhead parking
FISHUP is the Upper Fish Creek Trailhead parking
HRTBAR is the junction of SR38 and 1N05.
MNSHFT is Mine Shaft Saddle
SNGORG is Mount San Gorgonio.

Highlights of This Trail

The botanical highlights of this trail are:

The trail has five Asteraceae taxa with similar-looking flower heads with purple ligules; a key to separate them is given below.

Number of Unique Taxa On This Trail

The following histogram gives the number of trails in our database that contain each taxon on this trail. We had 77 trails in our database when this histogram was made; two of those trails, including this one, are in the Fish Creek Area, with two others in the San Bernardino Mountains. A number of "1" means the taxon has only been found on this trail among the trails in our database; numbers of "4" or smaller may indicate taxa found only in the San Bernardino Mountains.

Number of Trails
Containing A Taxon
Number Of Taxa
On This Trail
% of Taxa
On This Trail
Total Taxa147100%

Every taxon on the trail is included in the above table, although one of the taxa has not yet been keyed to a subspecies.

Fieldwork Dates and Summary of List Changes With Time

The following table gives the dates the trail was walked and taxa recorded. After each visit, the table gives the total number of taxa on the list and the breakdown of the taxa without positive identification. See Explanation of Plant Trail Guides to understand the symbols below.

Visit DateVisit ## taxa# "?"# "sp"# "~"# "ssp"

Only the first 0.24 miles of the trail, before the water crossing, was surveyed on 8/5/2003. Only the first 1.14 miles was surveyed on 5/27/04.

Botanical Trip Reports

27 May 2004
7 June 2004
11 June 2004
22 July 2004
10 August 2004

See also reports from the Upper Fish Creek Trail.

The Plant Guide

Version for printing, without lines and other text on this page: html (8 pages) or pdf Clickbook booklet (2 double-sided pages). (See printing instructions for an explanation of these options)

MileS#id?Common NameLatin Name#here#all
0.00   Trailhead at parking lot in front of kiosk / display board, elevation 7410 feet (2260 m). Trail is in sun for first 1/4 mile, then mostly in shade. (Coville's fleabane, Erigeron breweri var. covillei, is found in the parking area, but not so far on the trail).
0.00b1 threadleaf common rabbitbrush Chrysothamnus nauseosus ssp. consimilis20 / 514
0.00r2 hoary-aster Machaeranthera canescens var. canescens+50 / 93
0.00b3 Wright's buckwheat Eriogonum wrightii var. subscaposum40 / 611
0.00l4 California squirreltail Elymus elymoides ssp. californicus50 / 98
0.00r5 Nevin's bird's beak Cordylanthus nevinii30 / 55
0.00l6 yarrow Achillea millefolium20 / 411
0.00b7 Martin's paintbrush Castilleja applegatei ssp. martinii+30 / 414
0.00r  Information board
0.00r8 Davidson's lotus Lotus nevadensis var. davidsonii10 / 37
0.00r9 pinewoods rock-cress Arabis holboellii var. pinetorum99 / 92
0.00l10 *desert crested wheatgrass Agropyron desertorum3 / 13
0.00l11 matchweed Gutierrezia sarothrae20 / 14
0.00r12 spreading fleabane Erigeron divergens+50 / 93
0.00r13 California brome Bromus carinatus var. carinatus10 / 314
0.00l14 *downy brome Bromus tectorum99 / 919
0.00l15 Parish's buckwheat Eriogonum parishii20 / 33
0.00l16 groundsmoke Gayophytum diffusum ssp. parviflorum99 / 911
0.00l17 San Gabriel beardtongue Penstemon labrosus30 / 96
0.00r18 California black oak Quercus kelloggii1 / 111
0.00r19 green-leaf manzanita Arctostaphylos patula99 / 910
0.00r20 California-aster Lessingia filaginifolia var. filaginifolia30 / 541
0.00r21 Mojave linanthus Linanthus breviculus50 / 24
0.00l22 sulphur buckwheat Eriogonum umbellatum var. munzii30 / 49
0.00r23 plain mariposa lily Calochortus invenustus2 / 27
0.00r24 curl-leaf mountain-mahogany Cercocarpus ledifolius var. intermontanus5 / 38
0.00l  Jct. "scramble path" from parking lot.
0.00b25 Fendler's blue grass Poa fendleriana ssp. longiligula99 / 92
0.01l26 golden yarrow Eriophyllum confertiflorum var. confertiflorum30 / 546
0.01l27 dense false-gilia Allophyllum gilioides ssp. violaceum10 / 11
0.01l28 Johnston's knotweed Polygonum douglasii ssp. johnstonii10 / 11
0.01l29 needle & thread grass Hesperostipa comata ssp. comata5 / 11
0.01l30 Ross' sedge Carex rossii30 / 92
0.01l  Jct. trail from south end of parking lot.
0.01l31 one-sided bluegrass Poa secunda ssp. secunda30 / 521
0.02r32 Brewer's fleabane Erigeron breweri var. breweri30 / 51
0.02r33 purple-root cryptantha Cryptantha micrantha99 / 56
0.02r34 mountain mugwort Artemisia ludoviciana ssp. incompta99 / 95
0.02r35 granite prickly phlox Leptodactylon pungens30 / 54
0.03r36 Jeffrey pine Pinus jeffreyi50 / 919
0.03r37 forest goosefoot Chenopodium atrovirens99 / 93
0.04r38 canyon lotus Lotus argyraeus var. argyraeus4 / 21
0.04l39 narrowleaf monardella Monardella linoides ssp. stricta50 / 96
0.05r40 white fir Abies concolor99 / 918
0.05l41 goldenrod Solidago californica50 / 526
 r42 Modoc gilia Gilia modocensis5 / 13
0.09l43 canyon live oak Quercus chrysolepis3 / 219
0.09l44 mountain whitethorn Ceanothus cordulatus40 / 913
0.09l45 western wallflower Erysimum capitatum ssp. capitatum99 / 915
0.11l46 Parry's sunflower Hulsea vestita ssp. parryi40 / 72
0.11r47 wild tarragon Artemisia dracunculus50 / 521
0.11l48 splendid gilia Gilia splendens ssp. splendens50 / 26
0.12r49 purple nightshade Solanum xanti5 / 112
0.12r  (California false-indigo, Amorpha californica var. californica)
0.12l50 spear-leaved mountain dandelion Agoseris retrorsa5 / 212
0.13r51 Davidson's buckwheat Eriogonum davidsonii40 / 216
0.13l52 slender everlasting Gnaphalium canescens ssp. thermale20 / 32
0.14l53 southern montane grape lupine Lupinus excubitus var. austromontanus20 / 23
0.15l54 pine cryptantha Cryptantha simulans5 / 11
0.15b55 southern mountain woolly-star Eriastrum densifolium ssp. austromontanum+20 / 39
0.16l56 Parish's needlegrass Achnatherum parishii+10 / 25
0.16l  (bird's-foot fern, Pellaea mucronata var. mucronata)
0.17l57 slender bedstraw Galium angustifolium ssp. gracillimum+1 / 11
0.17l58 Wheeler's common madia Madia elegans ssp. wheeleri10 / 12
0.17l59 *yellow salsify Tragopogon dubius+10 / 25
0.17l60 incense-cedar Calocedrus decurrens4 / 311
0.22l61 California false-indigo Amorpha californica var. californica2 / 22
0.22b62 smoothleaf yerba santa Eriodictyon trichocalyx var. trichocalyx3 / 16
0.22c63 wild pepper-grass Lepidium virginicum var. pubescens10 / 35
0.22r64 woodland spurge Euphorbia palmeri10 / 32
0.27r65 interior rose Rosa woodsii var. ultramontana50 / 55
 l  (spineless horsebrush, Tetradymia canescens)
0.30   Trail turns right to cross stream; the rest of the trail is now almost entirely in shade.
0.30r  Sign: "Wilderness permit required before entry". (free; available from Mill Creek Forest Service Office)
0.31l66 mountain pink currant Ribes nevadense30 / 56
0.31l67 Richardson's geranium Geranium richardsonii40 / 53
0.31l68 mountain sweet-cicely Osmorhiza chilensis3 / 15
0.31l69 arroyo willow Salix lasiolepis30 / 528
0.31l70 hairy wood rush Luzula comosa5 / 21
0.31l71 Kentucky blue grass Poa pratensis ssp. agassizensis5 / 24
0.31l72 slender hairgrass Deschampsia elongata10 / 22
0.31l73 scarlet monkeyflower Mimulus cardinalis3 / 18
0.31l74 western columbine Aquilegia formosa20 / 55
0.31b75 willowherb Epilobium ciliatum ssp. ciliatum30 / 316
0.31l76 winter cress Barbarea orthoceras20 / 26
0.31l77 *perennial mouse-ear chickweed Cerastium fontanum ssp. vulgare20 / 22
0.31l78 cow parsnip Heracleum lanatum1 / 12
0.31l79 fragile sheath sedge Carex fracta5 / 16
0.31l80 Southern California draba Draba corrugata var. corrugata20 / 32
0.31l81 small white violet Viola macloskeyi5 / 11
0.31l82 rigid hedge-nettle Stachys ajugoides var. rigida3 / 13
0.31l83 stinging nettle Urtica dioica ssp. holosericea3 / 210
0.31l84 lemon lily Lilium parryi6 / 33
0.31l85 thyme-leaved speedwell Veronica serpyllifolia ssp. humifusa10 / 11
0.31r86 common monkeyflower Mimulus guttatus30 / 32
0.31r87 swamp sedge Carex senta99 / 92
0.31r88 field horsetail Equisetum arvense25 / 32
0.31r89 American speedwell Veronica americana10 / 12
0.31r90 Scouler's St. Johnswort Hypericum formosum var. scouleri10 / 13
0.31l  (yellow willow, Salix lutea; large-leaf avens, Geum macrophyllum)
0.31   Cross Fish Creek which has flowing water; elevation ~7280 feet (2220 m), the low point on the trail.
0.32r91 quaking aspen Populus tremuloides30 / 11
0.33l  Sign: "San Gorgonio Wilderness".
0.33l  (white bog orchid, Platanthera leucostachys)
0.34r  Jct. Aspen Grove Trail. Go left on Fish Creek Trail
0.34b92 smooth scouring rush Equisetum laevigatum99 / 93
0.34b93 Parish's snowberry Symphoricarpos rotundifolius var. parishii99 / 910
0.36l94 *prickly lettuce Lactuca serriola1 / 126
0.36r95 wax currant Ribes cereum var. cereum5 / 211
0.36l96 alkali western tansy-mustard Descurainia pinnata ssp. halictorum50 / 44
0.36r97 Fendler's meadow-rue Thalictrum fendleri var. fendleri10 / 43
0.37r98 creeping wild rye Leymus triticoides99 / 96
0.39l99 Utah service-berry Amelanchier utahensis2 / 27
0.39   Cross small dry drainage
0.41   Switchback left.
0.42   Trail curves right.
0.43r100 beaked penstemon Penstemon rostriflorus30 / 910
0.43r101 black cottonwood Populus balsamifera ssp. trichocarpa+99 / 96
 l102 3yellow willow tSalix lutea /
0.44l103 Grinnell's beardtongue Penstemon grinnellii var. grinnellii20 / 413
0.45   Trail curves right.
0.46l  Plants here have narrower leaves, almost like those of the former taxon of narrow-leaved black cottonwood, Populus trichocarpa var. ingrata.
0.46b104 mountain spray Holodiscus microphyllus var. microphyllus3 / 16
0.48l105 Lemmon's willow Salix lemmonii2 / 22
0.48r106 Anderson's lupine Lupinus andersonii+10 / 42
0.55l  Optional excursion to descend creek bank 15 steps to creek, which has the second occurrence of lemon lily, and three species also found later on the trail: horsetail, Equisetum hyemale ssp. affine, variegated clover,Trifolium variegatum phase 2, and cobwebby hedge-nettle, Stachys albens.
0.56r107 pine lousewort Pedicularis semibarbata20 / 99
0.57l  Jct. path to water. Stay on trail.
0.60l108 ranger's buttons Sphenosciadium capitellatum10 / 24
0.65r  Giant incense cedar (Calocedrus decurrens) with huge lightning scar. It is still alive despite having no visible bark!
0.69r109 Parish's bedstraw Galium parishii10 / 54
0.70l110 San Bernardino beardtongue Penstemon caesius10 / 93
0.70l111 Scouler's willow Salix scouleriana5 / 43
0.73b  (blue elderberry, Sambucus mexicana)
  112 scattered blazing star Mentzelia dispersa10 / 23
0.81l113 fir dwarf-mistletoe Arceuthobium abietinum / 2
0.82l114 blue elderberry Sambucus mexicana5 / 539
0.82   Cross dry drainage.
0.88b115 Parish's campion Silene parishii20 / 46
0.88r116 perennial rock-cress Arabis perennans5 / 12
0.94r117 ragged-leaf bahia Bahia dissecta2 / 11
0.94r  Sign high on tree: "No camping here".
0.99l118 bristly-leaved rock-cress Arabis rectissima var. rectissima+5 / 22
1.00   Cross dry drainage.
1.10b119 rosy everlasting, pussytoes Antennaria rosea ssp. confinis20 / 11
1.11r120 Nevada cinquefoil Potentilla glandulosa ssp. nevadensis20 / 39
1.14r121 horsetail Equisetum hyemale ssp. affine30 / 13
1.16   Cross small seasonally-moist drainage.
1.16l  (arrowhead butterweed, Senecio triangularis; sparse-flowered bog-orchid, Platanthera sparsiflora)
1.16l122 meadow starwort Stellaria longipes var. longipes3 / 11
1.16b123 giant red paintbrush Castilleja miniata ssp. miniata10 / 25
1.16r124 pearlwort Sagina saginoides /
1.16r  (little false-solomon's-seal, Smilacina stellata)
1.16r125 variegated clover Trifolium variegatum phase 2+50 / 22
1.16r126 long-anthered rush Juncus macrandrus20 / 12
1.17   Third location of lemon lily.
1.19r127 Parish's lupine Lupinus latifolius var. parishii6 / 13
1.19r128 fireweed Epilobium angustifolium ssp. circumvagum10 / 13
1.20r129 Parish's alumroot Heuchera alpestris1 / 12
1.20r130 larger mountain monkeyflower Mimulus tilingii /
1.20r131 musk monkeyflower Mimulus moschatus20 / 2
1.21 132 fir mistletoe Phoradendron pauciflorum1 / 19
1.21b133 Idaho bentgrass Agrostis idahoensis10 / 12
1.22r134 *common dandelion Taraxacum officinale25 / 24
1.22r  Gopher tunnel "casts" from gophers making tunnels in snow and then filling them with dirt.
 r135 San Bernardino rubber rabbitbrush Chrysothamnus nauseosus ssp. bernardinus /
1.23r136 spike bentgrass Agrostis exarata5 / 13
1.26l137 Mexican rush Juncus mexicanus30 / 510
1.28   Cross small drainage and enter Monkey Flower Flat.
1.28b138 fringed brome Bromus ciliatus5 / 13
1.28b139 Letterman's needlegrass Achnatherum lettermanii5 / 13
1.28r140 Big Bear Valley milk-vetch Astragalus lentiginosus var. sierrae5 / 11
1.29b141 Wheeler's cinquefoil Potentilla wheeleri20 / 12
1.29b142 mat muhly Muhlenbergia richardsonis2 / 12
1.30r  (pussy paws, Calyptridium monospermum)
1.30r143 Parry's sand cress Calyptridium parryi var. parryi20 / 23
1.33b144sspCalifornia evening-primrose Oenothera californica+10 / 12
1.33b145 fine-flower gilia Gilia leptantha ssp. leptantha20 / 21
1.37b146 spineless horsebrush Tetradymia canescens20 / 95
1.48l  Field of young pussypaws, Calyptridium monospermum.
1.48l147 pussy paws Calyptridium monospermum20 / 24
1.57   Cross Fish Creek.
1.59l  Jct. (old?) trail; continue straight.
1.59l  A mature short black cottonwood, 5 feet high, trunk 5 cm diameter, with lvs ~2 cm in width in 2002; 2.5-4.5 cm in width, most 3.0 cm, in 2003; and 2.0-3.1 cm in width, most 2.7-3.0 cm, in 2004.
1.61   Trail jags left.
1.64l  Field of mature pussypaws, Calyptridium monospermum.
1.66b148 Jepson's blue wildrye Elymus glaucus ssp. jepsonii10 / 19
1.66r149 cobwebby hedge-nettle Stachys albens3 / 14
1.66   Cross small drainage.
1.70b150 spreading dogbane Apocynum androsaemifolium+10 / 13
1.71   Trail jags left.
1.79   Switchback left.
1.80   Switchback right.
1.87l151 clustered fleabane Erigeron aphanactis var. congestus+6 / 21
1.93l  Jct. (old trail to Lower Fish Creek Meadow?)
2.00   End trail at jct. with Upper Fish Creek Trail, elevation 7980 feet (2430 m); see its plant trail guide to continue, or return the way you came.

Comments On Specific Species

Machaeranthera canescens var. canescens. Specimens from 9/27/02 had ray flowers that were not fertile - they had no styles, and their fruit was undeveloped. This made the keying to the variety impossible.

Specimens from 8/10/04 had fertile ray flowers with styles present, and hence keyed directly to var. canescens.

The JM description says that var. canescens can have ray flowers that are rarely reduced or 0.

Castilleja applegatei ssp. martinii. This taxon in the San Bernardino Mountains is different from the ones elsewhere!

Using Munz, the plants key to Castilleja martinii Abrams var. ewanii (Eastw.) Munz. Munz had three subspecies of C. martinii; this is the one found only in the San Bernardino Mountains.

The most striking distinction of var. ewanii is the leaf, which is linear and not wavy, unlike the lanceolate wavy leaves of var. martinii. There are also minor differences in the lengths of the galea (beak) and the depth of the calyx division. In addition to these distinctions listed in Munz, these plants are also much more glandular than the many specimens of var. martinii we have seen elsewhere.

In the JM, the three varieties of C. martinii were combined into the single subspecies of Castilleja applegatei.

Surprisingly, the name Castilleja martinii Abrams var. ewanii (Eastw.) Munz is listed in the JM as a synonym of C. angustifolia, which is non-glandular! In the JM entry for C. angustifolia, it says "earliest epithet in the widespread C. chromosa complex".

C. chromosa is in Munz, and is indeed non-glandular.

However, other sources, using Kartesz (1996), list Castilleja martinii var. ewanii as a synonym for Castilleja applegatei ssp. martinii.

Oddly, IPNI lists no synonyms between Castilleja martinii var. ewanii and Castilleja applegatei.

We are definitely confused about what has happened to the name of Castilleja martinii var. ewanii.

The story gets even more complicated. In 1980, Heckard defined the taxon C. montigena, and classified a voucher from this trail as that identification. In the JM, this taxon is called a hybrid of Castilleja applegatei ssp. martinii with C. angustifolia, which is listed as a judgement-reserved taxon under Castilleja applegatei ssp. martinii.

Since the JM name for these plants is Castilleja applegatei ssp. martinii, we've gone with that name. However, note that the JM description does not include or discuss the characteristics of Castilleja martinii var. ewanii mentioned above, instead leaving the reader to infer that those differences are due to its hybrid nature.

Erigeron divergens. There were no specimens of this species on the trail during our field work in 2002 and 2003! But in 2004, there were at least 50 plants spread along the first 0.20 miles of trail.

Although the JM considers this taxon to be an annual, and we have observed annuals of this species at lower elevations, Munz treats this as a biennial or nearly so. We suspect that the plants on this trail are almost entirely biennials. The severe drought in 2001-2002 killed all the biennial plants that germinated in spring 2001, producing no display in 2002. Plants that germinated in spring 2003 did not bloom in that year, resulting in no display in 2003. Finally, these plants produced their display in spring / summer 2004, surprising us greatly by their presence.

Eriastrum densifolium ssp. austromontanum. We treat ssp. elongatum as part of ssp. austromontanum, following blue Munz, since we generally find plants that have bracts that are 3-5 lobed, fitting ssp. elongatum, but have well over 15 flowers per head, fitting ssp. austromontanum. This is usually a symptom that the subspecies are not truly distinct.

Achnatherum parishii. This taxon may not be distinct from A. coronatum. See Achnatherum coronatum / A. parishii for a discussion of the characteristics of the specimens on this trail versus the characteristics of the two taxa given in the floras.

Galium angustifolium ssp. gracillimum. At first glance, these specimens seem identical with ssp. angustifolium. The difference is small, and comes down to this couplet in the JM:

60. Pl tall or low, ± stout, glabrous to white-hairy; lvs not deciduous; widespread in hills, mtns, but not in D ........ ssp. angustifolium
60'. Pl tall, very slender, gen glabrous; lvs deciduous; DMtns..... ssp. gracillimum

Note that for tall, glabrous plants, which fits most ssp. angustifolium, the only difference between the two subspecies is very slender vs. ± stout, and deciduous leaves or not. The leaves on these specimens are indeed deciduous, and the plants are on the slender side, so we have called them as ssp. gracillimum.

There are a handful more specimens off-trail nearby the one plant that is on the trail.

Tragopogon dubius. This non-native invasive species is only present in this portion of the trail. We pull up all plants when we see them, and take the flowers and seedheads back to our car and throw them in the trash. If everyone would do the same, we might be able to eliminate this before it becomes a serious pest, as it has elsewhere.

Populus trichocarpa sspp. trichocarpa and ingrata. Oddly, ssp. ingrata is possibly the second subspecies or variety that we have noticed that has gotten lost or been ignored in the JM. There are many specimens of this variety along the trail.

We were suspicious for some time that these specimens were normal Populus balsamifera ssp. trichocarpa that had simply been cut back, and whose new leaves were much narrower than the usual 3-7 cm quoted in the JM. Our suspicions were allayed when we found the specimen at mile 1.59, that clearly was very old and had not been cut back. Every leaf on that shrub had a width of ~2 cm in 2002, which is very different from the Populus balsamifera ssp. trichocarpa on the trail. Interestingly, the width of the leaves varied in subsequent years; they were 2.5-4.5 cm, most 3.0 cm, in 2003; and 2.0-3.1 cm in width, most 2.7-3.0 cm, in 2004.

Thus either this taxon needs to be restored to the JM, or else the leaf shape and width reported for Populus balsamifera ssp. trichocarpa needs to be extended.

Lupinus andersonii. The first specimen on the trail was too young to bloom in 2002, 2003 and 2004; the id comes from similar specimens that bloomed at mile 1.47 in 2003. We observed blooms from closer specimens in 2004, but did not record their location.

Arabis rectissima var. rectissima. All specimens on this trail and on the Upper Fish Creek Trail were infected with rust fungus, producing a large number of brown dots on almost every surface of the plants. This makes this Arabis easy to identify at any stage!

Potentilla glandulosa ssp. reflexa. All specimens on this trail clearly key to ssp. reflexa, having petals obovate; sepals > petals; st and pedicel hairs gen glandular (some non-glandular); lflet double-toothed, which is essentially the JM key. However, the petals are white, unlike the yellow petals of ssp. reflexa, and the inflorescence branch angles are 5-10°, unlike the gen 20-40° angles of ssp. reflexa. Both these characteristics fits ssp. nevadensis, so perhaps there is some intergradation occurring here.

Trifolium variegatum phase 2. According to a note in the JM, this taxon is commonly confused with T. wormskioldii, and we understand why. These plants look to be perennial, and the leaves and stems are almost the spitting image of the illustration in the JM. Using Munz, the only possible id is in fact T. wormskioldii, since apparently Munz describes only a subset of the T. variegatum population that does not fit these specimens.

However, they key perfectly to T. variegatum in the JM, since the involucres are only 1 cm wide, and there are 1-2 seeds per pod. Confirming the id, the inflorescence widths are 1.5-2.0 cm, and the JM notes that this can be a short-lived perennial. The inflorescence in fact looks much more like the JM illustration for T. variegatum than for T. wormskioldii.

In summary, there are no inconsistencies with a T. variegatum id for these plants, whereas there are five inconsistencies with a T. wormskioldii (the involucre is not gen 2-3 cm wide; the involucre is lobed; the seeds are not 2-6; the inflorescence width is not 2-3 cm; and the corolla tip is not white).

Oenothera californica. The subspecies here is not clear based solely on the taxonomy. The JM key is:

8. cauline lvs gen ± pinnately-lobed; pl ±grayish-green .... ssp. avita
8'. cauline lvs subentire to deeply wavy-dentate; pl green to slightly grayish .... ssp. californica

The leaves probably go with 8', since they are pinnatifid. But with the two weasel words in 8 (gen and ±), they could fit that, too. The leaves appear grayish-green, which would go with 8. However, based on the geography of the subspecies in the JM, the choice of 8 would lead to the wrong ssp, one found only in D in Southern California. However, Munz says of O. californica (presumably ssp. californica based on the geographic range): ashy with short appressed hairs, which fits our samples precisely.

Hence we have left the subspecies indeterminate.

Apocynum androsaemifolium. The specimens on this trail are significantly taller than the heights given in the floras. Tom measured heights of 2.7-5.2 dm for all the nine specimens in one area of the trail, compared to the 1.6-3 dm in the JM key.

Erigeron aphanactis var. congestus. The specimens on this trail have yellow ligules, unlike the orangish ligules for the Pebble Plains version. This taxon has a disjunct distribution in the San Bernardino Mountain, at the Pebble Plains and here in the San Gorgonio alpine zone, 6,000 - 11,000 feet elevation.

Aster separation.

Here is a key to separate the five similar Asteraceae on this trail with purple ligules:

1. phyllary tips appressed; lvs reduced upwards or not; annual to per ... Erigeron

2. phyllaries strongly graded; basal lvs 0, cauline gen evenly sized and spaced; infls arising near st tips; per
3. phyllaries glandular .... var. breweri
3'. phyllaries non-glandular ... var. covillei
2'. phyllaries roughly equal; cauline lvs reduced upwards; infls arising near mid-stem; ann to biennial E. divergens

1'. phyllary tips spreading to reflexed; lvs gen reduced upwards; per to subshrub
2. lvs green, linear, often toothed; resembling those of a penstemon in being rigidly spreading; gen per ..... Machaeranthera canescens
2'. lvs gray, "half-tomentose", usually ascending; subshrub ...... Lessingia filaginifolia var. filaginifolia

We thank Mike Crouse, James Dillane, Walt Fidler, and Dave Stith for help with the 2 July 2013 fieldwork.

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Updated 14 October 2013.